Feces (or faeces) are the solid or semisolid remains of the food that could not be digested in the small intestine. Bacteria in the large intestine further break down the material. Feces contain a relatively small amount of metabolic waste products such as bacterially altered bilirubin, and the dead epithelial cells from the lining of the gut. Feces are discharged through the anus or cloacaduring a process called defecation. Feces can be used as fertilizer or soil conditioner in agriculture. It can also be burned and used as a fuel source or dried and used as a construction material. Some medicinal uses have been found. In the case ofhuman feces, fecal transplants or fecal bacteriotherapy are in use. Urine and feces together are called excreta. The distinctive odor of feces is due to bacterialaction. Gut flora produces compounds such as indole, skatole, and thiols (sulfur-containing compounds), as well as the inorganic gas hydrogen sulfide. These are the same compounds that are responsible for the odor of flatulence. Consumption of foods prepared with spices may result in the spices being undigested and adding to the odor of feces. The perceived bad odor of feces has been hypothesized to be a deterrent for humans, as consuming or touching it may result in sickness or infection. Human perception of the odor may be contrasted by a non-human animal’s perception of it; for example, an animal who eats feces may be attracted to its odor. Feces are discharged through the anus or cloacaduring a process called defecation. This process requires pressures that may reach 100 mm Hg in humans and 450 mm Hg in penguins. The forces required to expel the feces are generated through muscular contractions and a build-up of gases inside the gut, prompting the sphincter to relieve the pressure on it and to release the feces. After an animal has digested eaten material, the remains of that material are discharged from its body as waste. Although it is lower in energy than the food from which it is derived, feces may retain a large amount of energy, often 50% of that of the original food. This means that of all food eaten, a significant amount of energy remains for the decomposers of ecosystems. Many organisms feed on feces, from bacteria to fungi to insects such as dung beetles, who can sense odors from long distances. Some may specialize in feces, while others may eat other foods. Feces serve not only as a basic food, but also as a supplement to the usual diet of some animals. This process is known as coprophagia, and occurs in various animal species such as young elephants eating the feces of their mothers to gain essential gut flora, or by other animals such as dogs, rabbits, and monkeys. Feces and urine, which reflect ultraviolet light, are important to raptors such as kestrels, who can see the near ultraviolet and thus find their prey by their middens and territorial markers. Seeds also may be found in feces. Animals who eat fruit are known as frugivores. An advantage for a plant in having fruit is that animals will eat the fruit and unknowingly disperse the seed in doing so. This mode of seed dispersal is highly successful, as seeds dispersed around the base of a plant are unlikely to succeed and often are subject to heavy predation. Provided the seed can withstand the pathway through the digestive system, it is not only likely to be far away from the parent plant, but is even provided with its own fertilizer. Organisms that subsist on dead organic matter or detritus are known as detritivores, and play an important role in ecosystems by recycling organic matter back into a simpler form that plants and other autotrophs may absorb once again. This cycling of matter is known as the biogeochemical cycle. To maintain nutrients in soil it is therefore important that feces return to the area from which they came, which is not always the case in human society where food may be transported from rural areas to urban populations and then feces disposed of into a river or sea. Depending on the individual and the circumstances, human beings may defecate several times a day, every day, or once every two or three days. The extensive hardening that interrupts this routine for several days or more is called constipation. The appearance of human fecal matter varies according to diet and health. Normally it is semisolid, with a mucus coating. A combination of bile and bilirubin, which comes from dead red blood cells, gives feces the typical brown color. After the meconium, the first stool expelled, a newborn’s feces contain only bile, which gives it a yellow-green color. Breast feeding babies expel soft, pale yellowish, and not quite malodorous matter; but once the baby begins to eat, and the body starts expelling bilirubin from dead red blood cells, its matter acquires the familiar brown color. At different times in their life, human beings will expel feces of different colors and textures. A stool that passes rapidly through the intestines will look greenish; lack of bilirubin will make the stool look like clay.